Beijing Key Laboratory of Development and Quality Control of Ornamental Crops, Department of Ornamental Horticulture and Landscape Architecture, China Agricultural University, Beijing 100193, China. Biology and Food Engineering College, Fuyang Normal University, Fuyang 236037, China. Biotechnology Institute, Fujian Academy of Agricultural Sciences, Fuzhou 350001, China. Department of Biological Sciences, University of Toronto, Toronto, ON M1C 1A4, Canada. Department of Cell and Systems Biology, University of Toronto, Toronto, ON M5S 3G3, Canada.